Bi Di Miss American Pie: Number Two Greek and Latin Roots!

A little while ago, I started a new series all about the Latin and Greek roots in the scientific names of different animals!  IN THE LAST POST, we examined animals with the Greek and Latin roots for "one," and today, we are going to kick it up a notch: exactly one notch, to be precise!  Today, we are going to examine the roots for the word "two!"  Let's begin with the cardinal (i.e. one, two three, etc.) form in Greek!  There are actually several roots that work here, but the one most commonly seen in binomial nomenclature is the root "di!"  Let's DIve right in!

• Our first "di" today is a small flying creature called Dimorphodon, a member of the extinct group of reptiles called pterosaurs.  The name "Dimorphodon" comes from three roots, "di," "morph," and "don."  "Di," of course, means "two."  In this context, the root word "morph" means "form."  In mythology and fantasty, a being that can take more than one form is often said to be able to morph their appearance.  Finally, the root word "don" is one of my favorites (and is used a whole lot in giving animals their scientific names): it means "tooth."  Altogether now: two-form tooth.  This name refers to the fact that Dimorphodon actually has two different types of teeth in its jaw.  For mammals, that's nothing special, but amongst reptiles, that is pretty rare!

• Dimetrodon-another animal with two kinds of teeth!  "Di" and "don" still mean the same thing as they did in Dimorphodon (above), but there is a new root in between: "metro."  For this root, think of the term "metric."  The name "Dimetrodon" actually means "two measures of teeth!"  Dimetrodon's two types of teeth would, in the groups that it is ancestral to, one day evolve to become the varied types of teeth that we see in the mouths of mammals!  Dimetrodon is more closely related to mammals than it is to any group of living reptile, and all of us mammals did evolve from a Dimetrodon-like ancestor!  So remember, if anyone ever tells you that Dimetrodon is a dinosaur, tell them that Dimetrodon actually lived around 40 million years before the first dinosaur ever walked the Earth!  That'll show them.

Dimetrodon (left) attacks the primitive amphibian Eryops.  Much like in the skull of Dimorphodon, you can clearly see the much larger teeth in the front of the skull and the smaller teeth in the back of the skull of Dimetrodon.

• Let's travel forward to the Late Cretaceous Period, time of Tyrannosaurus and Triceratops, to meet Didelphodon, a primitive mammal about the size of the living Virginia opossum!  As a matter of fact, it is from the opossum that Didelphodon gets its name: "Didelphodon" translates to "opossum tooth," as Didelphis is the genus name for the Virginia opossum and several related species of opossum!  In turn, "Didelphis" means "double womb," which presumably refers to the fact that the opossum, like all marsupials, has its internal reproductive tracts where the baby will develop for a bit, and its external pouch, where the baby will develop until full term.  

• A fourth animal with "two" and "tooth" in its name is Diprotodon!  The middle root, "pro," in this name means "forward," like the word "proceed."  So the name "Diprotodon" actually means "two forward teeth."  A quick examination of the skull of this massive mammal quickly reveals why!  Although it looks like it might be some sort of ungodly large rodent, Diprotodon is actually a hippopotamus-sized wombat, the largest marsupial known to have walked the Earth!

• Diceratops is a genus of ceratopsian dinosaur that is often considered to actually be a Triceratops.  The name, which means "two-horned face," was later discovered to already belong to a type of insect, and changed to the name Nedoceratops.  Some other paleontologists believe that Nedoceratops is really the same animal as Triceratops, but I don't really know enough about Nedoceratops to have an informed opinion on the matter.  However, paleontologist Jack Horner believes that Nedoceratops is an intermediate growth form between Triceratops and Torosaurus, and since I don't agree with his ideas of Triceratops ontogeny and that I think Triceratops and Torosaurus are definitely distinct dinosaurs, that leads me to suspect that Nedoceratops is more likely distinct, and certainly doesn't bridge the gap between Triceratops and Torosaurus.

• The name of Diplodocus, which means "double beam" originates from the two rows of chevron that are on the underside of the animal.  This was originally thought to be a feature unique to Diplodocus, a defining characteristic that would set it apart from other closely related sauropods.  Since Diplodocus was named by paleontologist Othniel Charles Marsh in the late 1800s, this feature has since been discovered on a number of other sauropods, including Barosaurus, also from the Morrison Formation, like Diplodocus.

• Dilophosaurus, one of the stars of the original Jurassic Park movie, gets its name from the two crests on its head.  Last time, we met Monolophosaurus, which means "single-crested lizard."  Therefore, Dilophosaurus means "two-crested lizard!"  

While it is the Greek cardinal root for "two" that is used most frequently in binomial nomenclature, it is the Latin root that is most often used for multiples (i.e. once, twice, thrice, etc.), the root "bi."  You can probably think of several words right off the top of your head that use this root!  In fact, the word "biped," used to describe creatures that walk on two feet (as opposed to, say, a quadruped), comes from the two roots "bi" and "ped," with "ped" meaning "foot" in Latin.  So literally, "biped" means "two feet!"  Let's look at a few more!

• Marshosaurus bicentesmus - A theropod dinosaur from the Morrison Formation (one who has received "Full-Post Status," as you can see by clicking HERE).  The exact relationships of Marshosaurus to other theropods isn't exactly clear, but some people think it might even be some sort of primitive coelurosaur, while others think it is more closely related to Megalosaurus and kin.  Regardless of its phylogenetic relationships, the name of Marshosaurus is quite exciting!  The genus name "Marshosaurus" honors the famous paleontologist Othniel Charles Marsh, who did a lot of work in the Morrison Formation.  The species name "bicentesmus" refers to the fact that the species was described in 1976, the bicentennial of the United States.  The bicentennial is, of course, a 200 year anniversary, and the "bi" in the name distinguishes a 200 year anniversary from a 100 year anniversary, or a centennial.

Here are two shots of a specimen of Marshosaurus that were on display in the lab at the Denver Museum of Nature and Science last year.  This first picture is the right maxilla, which would have made up part of the front section of the animals snout.

Here we have more of the same specimen, on display at the same time and the same place.  You can see several vertebrae and ribs in this shot.

• Baeolophus bicolor - This is the scientific name of the tufted titmouse, a small woodland bird native to much of the eastern half of the United States.  These little birds live in holes in trees that have been abandoned by woodpeckers, and are closely related to chickadees and, of course, the other titmice.  I couldn't figure out what the genus name "Baeolophus" means, but it is pretty apparent that the species name "bicolor" refers to the fact that this little bird is gray on its back, and white on its underside.  Some of the other species in the genus, such as the juniper titmouse (Baeolophus ridgwayi), are simply all gray.  

• Diceros bicornis - Here we have the scientific name of the black rhinoceros, a "Critically Endangered" species of African rhino.  According to some sources, the black rhino often prefers to eat Acacia leaves, presumably employing its prehensile upper lip to avoid the plants thorns.  The black rhino has pretty poor vision, with much better auditory and olfactory sensing capabilities.  Humans are easily the most dangerous threat to the black rhinoceros, with lions and the spotted hyena occasionally taking young black rhinos as prey, and even more rarely attacking adults.  The scientific name of the black rhinoceros literally means "two-horn two-horn."  As we have already established, both "di" and "bi" are roots that mean two.  Both "ceros" and "corn" are roots that refer to horns: think "Triceratops" for ceros (three-horned face), and "unicorn" for corn (one-horn).  

Works Cited:

Dinosaur Provincial Park

As we talked about IN A PREVIOUS POST, dinosaurs are pretty big in Canada.  A large number of dinosaur species have been discovered up there, and one of the best places to find dinosaurs is a place near Calgary in Alberta called Dinosaur Provincial Park.  As a matter of fact, Dinosaur Provincial Park is a UNESCO World Heritage Site, along with 981 other properties "which the World Heritage Committee considers as having outstanding universal value."  Other famous sites include the Great Barrier Reef in Australia; the Galápagos Islands; Stonehenge; the Grand Canyon; and "Memphis and its Necropolis," the site of the Great Pyramids and the Great Sphinx of Giza!  Just to name a few.  Do you get the idea, though?  Dinosaur Provincial Park is kind of a big deal!

But why?  What makes a bunch of badlands with some dinosaur bones in them so important to Canada, much less a committee dedicated to protecting such international treasures as the Great Barrier Reef and the Great Pyramids?  What makes Dinosaur Provincial Park so GREAT?  (Get it?  Nevermind, it wasn't that funny anyways.)  Here's what the UNESCO website has to say about the park:

The property is unmatched in terms of the number and variety of high quality specimens, over 60 of which represent more than 45 genera and 14 families of dinosaurs, which date back 75-77 million years. The park contains exceptional riparian habitat features as well as "badlands" of outstanding aesthetic value.
The committee also included two main criterion that show why the park is so important:

Criterion (vii): Dinosaur Provincial Park is an outstanding example of major geological processes and fluvial erosion patterns in semi-arid steppes. These "badlands" stretch along 24 kilometers of high quality and virtually undisturbed riparian habitat, presenting a landscape of stark, but exceptional natural beauty.


Criterion (viii): The property is outstanding in the number and variety of high quality specimens representing every known group of Cretaceous dinosaurs. The diversity affords excellent opportunities for paleontology that is both comparative and chronological. Over 300 specimens from the Oldman Formation in the park including more than 150 complete skeletons now reside in more than 30 major museums.

Wow. Well that's a pretty big deal!  According to the website, between 1979 and 1991, a grand total of around 23,347 fossils were collected, including an amazing 300 dinosaur skeletons!  As mentioned above, the dinosaur skeletons represent every known group of Cretaceous dinosaurs.  (I assume that they mean every group that is known to live in North America at the time.)  Not only does the sheer amount of fossils allow for a more complete view of an extinct ecosystem, new dinosaurs and other animals have been discovered there, as well as potential behavior that can be inferred from the fossils! 

During the Late Cretaceous North America was divided by the Western Interior Seaway, a shallow, continental sea.  (To learn more about the seaway, check out a recent post I did on it by clicking HERE).  In Dinosaur Provincial Park, you can find the remains of both ocean going animals and land dwellers, as well!  The park is, of course, famous for its dinosaurs (as you could probably tell from its name).  But many of the marine creatures entombed in the rocks there and in the surrounding area are pretty awesome, as well!  For example, Hybodus, an interesting shark!  

Before we get to the dinosaurs, let's check out a few other cool creatures found in the park!  One of these is a creature we mentioned in a previous post: the post entitled "There Be Dragons," all about the monitor lizards!  In the post, I had a picture of a prehistoric monitor lizard named Palaeosaniwa attacking a flock of Ornithomimus.  Well, both of these creatures have been found in the park!  Below is the picture, created by talented paleo-artist James Field!  You can check out his website HERE!

Many animals have been discovered in the park, including turtles, crocodilians, and a ton of plants, but the only other non-dinosaur we are going to look at for now is a small little primitive marsupial mammal called Eodelphis!  Eodelphis, whose name means "early opossum," is thought to have weighed a little over a pound which, astonishingly, means that it was one of the largest mammals of its time!  It is thought to be related to Didelphodon, another Mesozoic marsupial mammal, who we shall meet in an upcoming post!

Now for the dinosaurs!  I'm going to start with an animal called Centrosaurus.  A ceratopsian dinosaur (just like Triceratops), thousands of individuals specimens of Centrosaurus have been discovered in a massive bonebed that extends for hundreds of meters!  While paleontologists disagree as to exactly what killed all of these animals, and in such immense numbers, the prevailing theory is that this was a herd of animals that drowned while trying to cross a river.  The individuals that make up the herd vary widely in age, which is one of the lines of evidence which supports the herd idea.  This is important evidence for paleontologists, as it indicates that these animals lived in groups!  

Another dinosaur that is found in the park is the small pachycephalosaur called Stegoceras, who is not to be confused with the similarly named and much more famous Stegosaurus!

Here is another fun dinosaur, called Chirostenotes!  This oviraptorosaur was first found in the park, and is definitely quite funky looking!

Dromaeosaurus, a distant cousin of Chirostenotes and a closer relative of the famous Velociraptor, was also first discovered in the park!

Just as Dromaeosaurus has been pushed from the limelight by Velociraptor, so too has Daspletosaurus been pushed by Tyrannosaurus!  Daspletosaurus is a tyrannosaur as well, and was first discovered in (surprise surprise) Dinosaur Provincial Park!  Two more Dinosaur Provinicial Park natives (and firsts) are Euoplocephalus, one of the tank-like ankylosaurs, and Parasaurolophus, a hadrosaur or duck-billed dinosaur!

I can't WAIT to visit the park one day!  In the meantime, HERE is a link to the park's website so you, too can plan your visit!

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